The importance of species-specific swim speeds for the formation of multi-species fish shoals in a Canadian freshwater system was investigated. Swim speed was positively correlated with total length and differed between species but was unrelated to shoal size.
Krause, J., Ward, A. J. W., Jackson, A. L., Ruxton, G. D., James, R., & Currie, S. (2005). The influence of differential swimming speeds on composition of multi-species fish shoals. Journal of Fish Biology, 67(3), 866-872. https://doi.org/10.1111/j.0022-1112.2005.00768.x